The forgotten Spiders of Mirkwood
In the Hobbit they were quite a big theme, even appearing on Thorin's map ('South lies Mirkwood the Great ~ there are Spiders); in LoTR, UT and the Appendixes they aren't even given a cursory reference.
A forgetful mistake, perhaps. It doesn't really matter much, but it's just one of those things that was never explained.
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