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Fatty Lumpkin-> Merry's ponies-> Nazgul's horses-> Fell beasts-> Gwaihir (and other eagles)-> Chief of the Wargs
Of course I'm only assuming that the different steeds of the Ringwraiths were connected, maybe being trained at the same time and place or something of the sort.
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'They say that the One will himself enter into Arda, and heal Men and all the Marring from the beginning to the end."
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