This is how I interpreted it: (be patient, it gets confusing.)
The Sun and Moon were created from the last fruit and the last flower of the Two Trees. The fruit from the Golden Tree Laurelin was placed in the sky by the Valar Varda, and was guided through the sky by the female Maier Arien.
The last flower of the White Tree Telperion was also placed in the sky by Varda, and was guided through the sky by the male Maier Tilion.
So my thought was that through the ages, the actual Sun and Moon assumed the female and male aspect of the Maier who guided them, at least in the minds of the people of Middle Earth.
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